The latest find of spilapterids: a new species Dunbaria elkunensis sp. Nov. (Insecta: Palaeodictyoptera: Spilapteridae) from the Middle Permian Golyusherma locality in Udmurtia, Russia

Cover Page

Cite item

Full Text

Open Access Open Access
Restricted Access Access granted
Restricted Access Subscription Access

Abstract

Based on the forewing found in the Middle Permian deposits of Golyusherma (Takhtashur) site, Udmurtia, a new paleodictyopteran species of Dunbaria elkunensis sp. nov. from the family Spilapteridae is described. The new species is the latest discovery of Spilapteridae, making this family the longest-lived among Palaeodictyoptera, the oldest representative of which is described from the Lower Carboniferous of Germany.

Full Text

Restricted Access

About the authors

N. D. Sinitshenkova

Borissiak Paleontological Institute, Russian Academy of Sciences

Author for correspondence.
Email: nina_sin@mail.ru
Russian Federation, Moscow, 117647

References

  1. Голубев В.К. Местонахождение нижнеказанских тетрапод Голюшерма (Удмуртия) // Бюлл. Моск. об-ва испыт. прир. Отд. геол. 1992. Т. 67. Вып. 3. С. 131.
  2. Голубев В.К. Событийная корреляция морских отложений казанского яруса стратотипической области // Стратигр. Геол. корреляция. 2001. Т. 9. № 5. С. 40–58.
  3. Горохов А.В. Пермские прямокрылые инфраотряда Oedischiidea (Orthoptera, Ensifera) // Палеонтол. журн. 1987. № 1. С. 62–75.
  4. Ефремов И.А., Вьюшков Б.П. Каталог местонахождений пермских и триасовых наземных позвоночных на территории СССР // Тр. Палеонтол. ин-та АН СССР. 1955. Т. 46. 186 с.
  5. Залесский М.Д. О новых насекомых из пермских отложений бассейнов рек Камы, Вятки и Белой // Тр. Об-ва естествоиспыт. при Казанском ун-те. 1929. Т. 52. Вып. 1. 30 с.
  6. Мартынов А.В. Пермские ископаемые насекомые Чекарды // Тр. Палеонтол. ин-та АН СССР. 1940. Т. 11. Вып. 1. 62 с.
  7. Шаров А.Г., Синиченкова Н.Д. Новые Palaeodictyoptera с территории СССР // Палеонтол. журн. 1977. № 1. С. 48–63.
  8. Beckemeyer R.J., Byers G.W. Forewing morphology of Dunbaria fasciipennis Tillyard (Palaeodictyoptera: Spilapteridae), with notes on a specimen from the University of Kansas Natural History Museum // J. Kansas Entomol. Soc. 2001. V. 74. P. 221–230.
  9. Béthoux O. Cladotypic taxonomy applied: titanopterans are orthopterans // Arthropod Syst. & Phylog. 2007. V. 65. № 2. P. 135–156.
  10. Béthoux O. King crickets, raspy crickets and weta, their wings, their fossil relatives // J. Orthoptera Res. 2012. V. 21. № 2. P. 179–225.
  11. Béthoux O., Nel A., Schneider J., Gang G. Lodetiella magnifica nov. gen. and nov. sp. (Insecta: Palaeodictyoptera; Permian), an extreme situation in wing morphology of palaeopterous insects // Geobios. 2007. V. 40. № 2. P. 181–189.
  12. Béthoux O., Voigt S., Schneider J.W. A Triassic palaeodictyopteran from Kyrgyzstan // Palaeodiversity. 2010. V. 3. P. 9–13.
  13. Brauckmann C., Gröning E. A new species of Homaloneura (Palaeodictyoptera: Spilapteridae) from the Namurian (Upper Carboniferous) of Hagen-Vorhalle (Germany) // Entomol. Gener. 1998. V. 23. № 1/2. P. 77–84.
  14. Brauckmann C., Herd K.J. Insekten-Fund aus dem Westfalium D (Ober-Karbon des Piesberges bei Osnabrück (Deutschland). Teil 1: Palaeoptera // Osnabrücker Naturwiss. Mitt. 2002. Bd 28. S. 27–69.
  15. Brauckmann C., Schneider J.W. A Lower Carboniferous insect from the Bitterfeld/Delitzsch area (Pterygota, Arnsbergian, Germany) // N. Jb. Geol. Paläontol. Monatsch. 1996. H. 1. S. 17–30.
  16. Carpenter F.M. Studies on North American Carboniferous insects. 3. A spilapterid from the vicinity of Mazon Creek, Illinois (Palaeodictyoptera) // Psyche. 1964. V. 71. № 3. P. 117–124.
  17. Handlirsch A. Die fossilen Insekten und die Phylogenie der rezenten Formen // Ein Handbuch für Paläontologen und Zoologen / Ed. Engelmann W. Leipzig: Engelmann, 1906–1908. 1430 s.
  18. Kukalová J. New Palaeodictyoptera (Insecta) of the Carboniferous and Permian of Czechoslovakia // Sborn. Ústředn. ústavu geol. Paleontologie. 1958. V. 25. P. 239–251.
  19. Kukalová J. Revisional study of the order Palaeodictyoptera in the Upper Carboniferous shales of Commentry, France, Part 1 // Psyche. 1969a. V. 76. № 2. P. 163–215.
  20. Kukalová J. Revisional study of the order Palaeodictyoptera in the Upper Carboniferous shales of Commentry, France, Part II // Psyche. 1969b. V. 76. № 4. P. 439–486.
  21. Kukalová J. Revisional study of the order Palaeodictyoptera in the Upper Carboniferous shales of Commentry, France, Part III // Psyche. 1970. V. 77. № 1. P. 1–44.
  22. Kukalová-Peck J. The structure of Dunbaria (Palaeodictyoptera) // Psyche. 1971. V. 78. № 4. P. 306–318.
  23. Laurentiaux D. Description de trois nouveaux Paléodictyoptères du terrain houiller du Nord de la France // Ann. Soc. Géol. Nord. 1949. V. 69. P. 206–223.
  24. Laurentiaux D., Teixeira C. Um novo género de insecto paleodictióptero do Carbónico continental do Baixo-douro (Portugal) // Bol. Soc. Geol. Portugal. 1958. V. 12. № 3. P. 41–49.
  25. Li Y., Ren D., Pecharová M., Prokop J. A new palaeodictyopterid (Insecta: Palaeodictyoptera: Spilapteridae) from the Upper Carboniferous of China supports a close relationship between insect faunas of Quilianshian (northern China) and Laurussia // Alcheringa. 2013. V. 37. P. 1–9.
  26. Liu X., Béthoux O., Yin X.C., Ren D. The smallest Palaeodictyoptera (Insecta) discovered at Xiaheyan (Late Carboniferous, China) // C. R. Palevol. 2015. V. 14. P. 346–352.
  27. Prokop J., Pecharová M., Jarzembowski E.A., Ross A.J. New Palaeodictyopterans from the Late Carboniferous of the UK (Insecta: Palaeodeictyoptera) // Earth and Environ. Trans. Roy. Soc. Edinburgh. 2017. V. 107. Iss. 2-3. P. 99–107. https://doi.org/10.1017/S1755691017000408
  28. Prokop J., Nel A., Engel M.S. et al. New Carboniferous fossils of Spilapteridae enlighten postembryonic wing development in Palaeodictyoptera // Syst. Entomol. 2015. V. 41. № 1. P. 178–190. https://doi.org/10.1111/syen.12148
  29. Prokop J., Ren D. New significant fossil insects from the Upper Carboniferous of Ningxia in northern China (Palaeodictyoptera, Archaeorthoptera) // European J. Entomol. 2007. V. 104. P. 267–275.
  30. Prokop J., Roques P., Nel A. New non-holometabolous insects from the Pas-de-Calaes, France (Insects: ’Exopterygota’) // Alcheringa. 2014. V. 38. P. 155–169. https://doi.org/10.1080/03115518.2014.848620
  31. Shcherbakov D.E. The alleged Triassic palaeodictyopteran is a member of Titanoptera // Zootaxa. 2011. V. 3044. P. 65–68.
  32. Sinitshenkova N.D. Chapter 2.2.1.2.3.1. Order Dictyoneurida Handlisch, 1906 (= Palaeodictyoptera Goldenberg, 1854) // History of Insects / Eds. Rasnitsyn A.P., Quicke D.L.J. Dordrecht, Boston, L.: Kluwer Acad. Publ., 2002. P. 116–120.
  33. Szwedo J., Grzywocz A., Grzywocz J. First Spilapteridae (Palaeodictyoptera) from Namurian b of Bielszowice coal mine, Upper Silesia // Fossil Record in Resins and Sediments. 25th Anniversary of Museum of Amber Inclusions. University Gdańsk 23rd-26th May 2023. Book of Abstracts. 2023. P. 106.
  34. Tillyard R.J. Kansas Permian insects. Part 1. The geologic occurrence and the environment of the insects with description of a new palaeodictyopterid // Amer. J. Sci. 1924. V. 5. № 7. P. 203–208.
  35. Tillyard R.J. Kansas Permian insects. Part 4. The order Palaeodictyoptera // Amer. J. Sci. 1925. V. 9. № 52. P. 328–335.
  36. Zalessky G. Sur un représentant d’un nouveau ordre des Hemiodonates de Permien de l’Oural // Bull. Soc. Natur. Moscou. Ser. Biol. 1946. V. 51. № 4–5. P. 63–70.

Supplementary files

Supplementary Files
Action
1. JATS XML
Download
2. Fig. 1. Dunbaria elkunensis sp. nov., holotype PIN, No. 5909/1, forewing without basal part; Udmurtia, Golyusherma locality; middle Permian, Kazanian stage. Scale bar is 5 mm.

Download (129KB)
Indexing metadata
3. Table III

Download (3MB)
Indexing metadata
4. Table IV

Download (3MB)
Indexing metadata

Copyright (c) 2025 Russian Academy of Sciences